Doña, J., Serrano, D., Mironov, S., Montesinos‐Navarro, A., Jovani, R.  (2018) Unexpected bird‐feather mite associations revealed by DNA metabarcoding uncovers a dynamic ecoevolutionary scenario. Molecular Ecology, DOI


Vizcaíno, A., Doña, J., Vierna, J., Marí-Mena, N., Esteban, R., Mironov, S., Urien, C., Serrano, D., Jovani, R.  (2018) Enabling large-scale feather mite studies: an Illumina DNA metabarcoding pipeline. Experimental and Applied Acarology, DOI


de Rojas, M., Doña J., Jovani, R., Dimov, I., Zurita, A., Callejón, R., Rodríguez-Plá, M.  (2018) Evidence of cryptic species in the genus Tinaminyssus (Acari: Rhinonyssidae) based on morphometrical and molecular data. Experimental and Applied Acarology, DOI


Esteban, R., Doña, J., Vierna, J., Vizcaíno, A., Serrano, D., Jovani, R. (2018) The complete mitochondrial genome of the feather mite Trouessartia rubecula Jablonska, 1968 (Astigmata: Analgoidea: Trouessartiidae). Mitochondrial DNA Part B,  DOI

Doña J., Proctor, H., Serrano, D., Johnson, KP., Oddy-van Oploo, A., Ascunce, MS., Huguet-Tapia, JC., Jovani, R (2018) Feather mites play a role in cleaning host feathers: New insights from DNA metabarcoding and microscopy. Molecular Ecology, DOI

Doña J., Proctor, H., Mironov, S., Serrano, D., Jovani, R (2017) Host specificity, infrequent major host switching and the diversification of highly host-specific symbionts: The case of vane-dwelling feather mites. Global Ecology and Biogeography, 27, 188-198, DOI

Carlson CJ., Burgio, KR., Dougherty, ER., Phillips, AJ., Bueno, VM., Clements, CF., Castaldo, G., Dallas, TA., Cizauskas, CA., Cumming GS., Doña,
., Harris, ND., Jovani, R ., Mironov, S., Muellerklein, OC., Proctor, HC., Getz, WM (2017) Parasite biodiversity faces extinction and redistribution in a changing climate. Science Advances, e1602422. DOI

Edelaar, P*., Jovani, R* ., Gomez-Mestre, I (2017) Should I Change or Should I Go? Phenotypic Plasticity and Matching Habitat Choice in the Adaptation to Environmental Heterogeneity. American Naturalist, 00: 000-000, pdfDOI (*Authors contributed equally to this work)

Doña, J., Sweet, AD., Johnson, KP., Mironov, SV., Serrano, D., Jovani, R (2017) Cophylogenetic analyses reveal extensive host-shift speciation in a highly specialized and host-specific symbiont system. Molecular Phylogenetics and Evolution, 00: 000-000, DOI

Vierna, J*.,  Doña, J*., Vizcaíno, A., Serrano, D., Jovani, R. (2017) PCR cycles above routine numbers do not compromise high-throughput DNA barcoding results. Genome. In press. DOI
(*Authors contributed equally to this work)

Jovani, R., Doña, J., Labrador, M.M., and Serrano, D. (2017) Opening the doors of parasitology journals to other symbionts. Trends in Parasitology, 33: 578-579, DOI

Doña, J., Potti, J., De la Hera, I., Blanco,G., Frias, and Jovani, R. (2017). Vertical transmission in feather mites: insights into its adaptive value. Ecological entomology, 42: 492-499, DOI

Pérez-Rodríguez, L., Jovani, R. & Stevens,M. (2017). Shape matters: animal colour patterns as signals of individual quality. Proc. R. Soc. B., 284: 20162446. DOI

Doña, J.*, Proctor, H.*, Mironov, S.*, Serrano, D., and Jovani, R. (2016). Global associations between birds and vane-dwelling feather mites. Ecology, 97: 3242. DOI: 10.1002/ecy.1528. (*Authors contributed equally to this work). pdf and dataset. **Please, read the propietary restrictions in the metadata before using for research purposes.

Jovani, R and Rohwer, S. (2016). Fault bars in bird feathers: mechanisms, and ecological and evolutionary causes and consequences. Biological reviews, 000-000. doi: 10.1111/brv.12273 pdf

Doña, J.*, Moreno-García, M.*, Criscione, CD., Serrano, D and Jovani, R. (2015). Species mtDNA genetic diversity explained by infrapopulation size in a host-symbiont system. Ecology and Evolution, 5:5801-5809. pdf (*Authors contributed equally to this work)

Doña, J.*, Ruiz-Ruano, FJ.*, Jovani, R. (2015). DNA barcoding of Iberian Peninsula and North Africa Tawny Owls Strix aluco suggests the Strait of Gibraltar as an important barrier for phylogeography. Mitochondrial DNA, 27(6):4475-4478 pdf (*Authors contributed equally to this work)

Jovani, R ., Lascelles, B., Garamszegi, L., Mavor, R., Thaxter, CB., and Oro, D. (2016). Colony size and foraging range in seabirds. Oikos, 125:968-974 pdf


Diaz‐Real, J., Serrano, D., Piriz, A., & Jovani, R. (2015). NGS metabarcoding proves successful for quantitative assessment of symbiont abundance: the case of feather mites on birds. Experimental and Applied Acarology,67(2):209-18. pdf

Doña, J., Diaz‐Real, J., Mironov, S., Bazaga, P., Serrano, D., & Jovani, R. (2015). Barcoding Feather Mites. The Barcode Bulletin. 6,2 pdf

Mironov, S., Doña, J., & Jovani, R. (2015) A new feather mite of the genus Dolichodectes (Astigmata:Proctophyllodidae) from Hippolais polyglotta (Passeriformes: Acrocephalidae) in Spain. Folia parasitologica, 62:032. DOI PDF

Doña, J., Diaz‐Real, J., Mironov, S., Bazaga, P., Serrano, D., & Jovani, R. (2015). DNA barcoding and mini‐barcoding as a powerful tool for feather mite studies. Molecular Ecology Resources 15: 1216-1225 DOI pdf

Sanz-Aguilar, A.*, Jovani, R.*, Melián, C. J., Pradel, R., & Tella, J. (2015). Multievent capture-recapture analysis reveals individual foraging specialisation in a generalist species. Ecology 96: 1650-1660 DOI pdf (*Authors contributed equally to this work)

Diaz-Real, J., Serrano, D., Pérez-Tris, J., Fernández-González, S., Bermejo, A., Calleja, J. A., … & Jovani, R. (2014). Repeatability of Feather Mite Prevalence and Intensity in Passerine Birds. PLoS ONE, 9(9), e107341. DOI pdf

Meléndez. L., Laiolo, P., Mironov, S., García, M., Magaña, O., & Jovani, R. (2014) Climate-driven variation in the intensity of a host-symbiont animal interaction along a broad elevation gradient. PLoS ONE 9(7): e101942. DOI pdf

sky networks

Cortés-Avizanda, A.*, Jovani, R.*, Donázar, J.A. & Grimm, V. (2014). Bird sky networks: how do avian scavengers use social information to find carrion? Ecology 95:1799–1808. DOI pdf, Suppl. Mat., NetLogo model (*Authors contributed equally to this work)

fault bars bacteria

Jovani, R., Montalvo, T., Savaté, S. (2014) Fault bars and bacterial infection. Journal of Ornithol 155, 3, DOI  pdf


Gomez-Mestre, I. & Jovani, R. (2013) A heuristic model on the role of plasticity in adaptive evolution: plasticity increases adaptation, population viability and genetic variation. Proceedings of the Royal Society B 280: 20131869

Jovani, R., Pérez-Rodríguez, L., Mougeot, F. (2013) Fractal geometry for animal biometrics: a response to Kühl and Burghardt. Trends in Ecology & Evolution 28: 499-500. doi:10.1016/j.tree.2013.06.004


Pérez-Rodríguez, L., Jovani, R., Mougeot, F. (2013) Fractal geometry of a complex plumage trait reveals bird’s quality. Proceedings of the Royal Society B 280: 20122783. doi:10.1098/rspb.2012.2783

Cortés-Avizanda, A., Jovani, R., Carrete, M. & Donázar, J.A. (2012) Resource unpredictability promotes species diversity and coexistence in an avian scavenger guild: a field experiment. Ecology 93: 2570-2579.

Canal, D. Jovani, R. & Potti, J. (2012) Male decisions or female accessibility? Spatiotemporal patterns of extra pair paternity in a songbird. Behavioral Ecology 23: 1146-1153. doi:10.1093/beheco/ars090

Jovani, R., Schielzeth, H., Mavor, R. & Oro, D. (2012). Specificity of grouping behavior: comparing colony sizes for the same seabird species in distant populations. Journal of Avian Biology 43: 397-402. doi: 10.1111/j.1600-048X.2012.05643.x

Baos, R., Jovani, R., Serrano, D., Tella, J.L. & Hiraldo, F. (2012) Developmental exposure to a toxic spill compromises long-term reproductive performance in a wild, long-lived bird: the white stork (Ciconia ciconia). PLoS ONE 7(4): e34716. doi:10.1371/journal.pone.0034716

Jovani, R. & Díaz-Real, J. (2012) Fault bars timing and duration: the power of studying feather fault bars and growth bands together. Journal of Avian Biology 43: 97-101. doi: 10.1111/j.1600-048X.2012.05583.x

Galván, I., Aguilera, E., Atiénzar, F., Barba, E., Blanco, G., Cantó, J.L. Cortés, V., Frías, O., Kovács, I., Meléndez, L., Møller, A.P., Monrós, J.S., Pap, P.L., Piculo, R., Senar, J.C., Serrano, D., Tella, J.L., Vágási, C.I., Vögeli, M. & Jovani, R. (2012) Feather mites (Acari: Astigmata) and body condition of their avian hosts: a large correlative study. Journal of Avian Biology 43: 273-279. doi: 10.1111/j.1600-048X.2012.05686.x

Beauchamp, G., Alexander, P. & Jovani, R. (2012) Consistent waves of collective vigilance in groups using public information about predation risk. Behavioral Ecology 23: 368-374.  doi:10.1093/beheco/arr194

Jovani, R., Avilés, J.M., & Rodríguez-Sanchez, F. (2012) Age-related sexual plumage dimorphism and badge framing in the European Robin Erithacus rubecula. Ibis 154: 147-154. doi: 10.1111/j.1474-919X.2011.01187.x

Canal, D. Jovani, R., & Potti, J. (2012) Multiple mating opportunities boost protandry in a pied flycatcher population. Behavioral Ecology and Sociobiology 66: 67-76. doi:10.1007/s00265-011-1253-8

Jovani, R., & Mavor, R. (2011) Group size versus individual group size frequency distributions: a nontrivial distinction. Animal Behaviour 82: 1027-1036. doi:10.1016/j.anbehav.2011.07.037

Galarza, J.A., Jovani, R., Cerdá, X., Rico, C., Barrosoa, A., & Boulay, R. (2012) Frequent colony relocations do not result in effective dispersal in the Gypsy ant Aphaenogaster senilis. Oikos 121: 605–613. doi: 10.1111/j.1600-0706.2011.19859.x

Jovani, R., Blas, J. Navarro, C. & Mougeot, F. (2011) Feather growth bands and photoperiod. Journal of Avian Biology 42: 1-4. doi: 10.1111/j.1600-048X.2010.05175.x

Jovani, R., Blas, J., Stoffel, M.J. Bortolotti, L.E. & Bortolotti, G.R. (2010) Fault bars and the risk of feather damage in cranes. Journal of Zoology 281: 94-98. doi:10.1111/j.1469-7998.2009.00686.x

Herrera, C.M. & Jovani, R. (2010) Lognormal distribution of individual lifetime fecundity: insights from a 23-year study. Ecology 91:422-430. doi:10.1890/09-0849.1

Jovani, R., Mavor, R. & Oro, D. (2008) Hidden patterns of colony size variation in seabirds: a logarithmic point of view. Oikos 117: 1774-1781. doi: 10.1111/j.1600-0706.2008.17065.x

Jovani, R. & Grimm, V. (2008) Breeding synchrony in colonial birds: from local stress to global harmony. Proceedings of the Royal Society of London B 275: 1557–1563. doi:10.1098/rspb.2008.0125

Jovani, R., Serrano, D., Ursúa, E. & Tella, J.L. (2008) Truncated power laws reveal a link between low-level behavioral processes and grouping patterns in a colonial bird. PloS ONE 3(4): e1992. doi:10.1371/journal.pone.0001992.

Jovani, R. & Fortuna, M.A. (2007) The shape of the past in the World Wide Web: scale-free patterns and dynamics. Physica A 385: 683–688. doi:10.1016/j.physa.2007.07.023

Jovani, R. & Tella, J.L. (2007) Fractal bird nest distribution produces power law colony sizes Proceedings of the Royal Society of London B 274: 2465-2469. doi:10.1098/rspb.2007.0527

Laiolo, P. & Jovani, R. (2007) The emergence of the animal culture conservation. Trends in Ecology & Evolution 22: 5. doi:10.1016/j.tree.2006.10.007

Smits, J. E., Bortolotti, G.R., Baos, R., Jovani, R., Tella, J.L., & Hoffmann, W.E. (2007). Disrupted bone metabolism in contaminant-exposed white storks (Ciconia ciconia) in southwester Spain. Environmental Pollution 145: 538-544. doi:10.1016/j.envpol.2006.04.032

Pueyo, S. & Jovani, R. (2006) Comment on “A keystone mutualism drives pattern in a power function”. Science 313: 1739c. doi: 10.1126/science.1129595

Baos, R., Jovani, R., Pastor, N., Tella, J.L., Jiménez, B., Gómez, G., González, M.J., & Hiraldo, F. (2006) Evaluation of genotoxic effects of heavy metals and arsenic in wild nestling white sortks (Ciconia ciconia) and black kites (Milvus migrans) from Southwestern Spain after a mining accident. Environmental Toxicology and Chemistry 25: 2794-2803. doi: 10.1897/05-570R.1

Jovani, R., Serrano, D., Frías, O. & Blanco, G. (2006) Shift in feather mite distribution during the molt of passerines: the case of barn swallows (Hirundo rustica). Canadian Journal of Zoology 84: 729-735. doi:10.1139/Z06-042

Jovani, R. & Tella, J.L. (2006) Parasite prevalence and sample size: misconceptions and solutions. Trends in Parasitology 22: 214-218. doi:10.1016/

Sarasola, J.H. y Jovani, R. (2006) Risk of feather damage explains fault bar occurrence in a migrant hawk, the Swainson’s hawk Buteo swainsoni. Journal of Avian Biology 37: 29-35. doi: 10.1111/j.2006.0908-8857.03693.x

Baos, R., Jovani, R., Forero, M.G., Tella, J.L., Gómez, G., Jiménez, B., González, M.J., & Hiraldo, F. (2006) Relationships between T-cell-mediated immune response and Pb, Zn, Cu, Cd, and As concentrations in blood of nestling White storks (Ciconia ciconia) and Black kites (Milvus migrans) from Doñana (Southwestern Spain) after the Aznalcóllar toxic spill. Environmental Toxicology and Chemistry 25:1153-1159. doi:10.1897/05-395R.1

Jovani, R. & Tella, J.L. (2005) Wear and opening as sources of band loss in the White Storks. Waterbirds 28(4): 426-429. doi:;2

Jovani, R. & Sol, D. (2005) How predictable is the abundance of double gametocyte infections?. Parasitology Research 97:84-86. doi: 10.1007/s00436-005-1405-8

Serrano, D. & Jovani, R. (2005) Adaptive fault bar distribution in a long-distance migratory, aerial forager passerine? Biological Journal of the Linnaean Society 85:455-461. doi: 10.1111/j.1095-8312.2005.00509.x

Jovani, R., Tella, J.L., Blanco, G. & Bertellotti, M. (2004) Variable inter-annual relationships between T-cell mediated immunity and individual traits in White Storks. Ardeola 51: 357-364.

Jovani, R., Amo, L., Arriero, E., Krone, O. Marzal, A., Shurunlinkov, P., Tomás, G., Sol, D., Hagen, J., López, P., Martín, J., Navarro, C., & Torres, J. (2004) Double gametocyte infections in Apicomplexan parasites of birds and reptiles. Parasitology Research 94: 155-157. doi: 10.1007/s00436-004-1186-5

Jovani, R. & Tella, J.L. (2004) Age-related environmental sensitivity and weather mediated nestling mortality in white storks (Ciconia ciconia). Ecography 27: 611-618. doi: 10.1111/j.0906-7590.2004.03925.x

Jovani, R. & Blas, J. (2004). Adaptive allocation of stress-induced deformities on bird feathers. Journal of Evolutionary Biology 17: 294-301. doi: 10.1111/j.1420-9101.2003.00680.x pdf

Pastor, N., Baos, R., López-Lázaro, M., Jovani, R., Tella, J.L., Hajji, N., Hiraldo, F., & Cortés, F. (2004) A 4-year follow-up analysis of genotoxic damage in birds of the Doñana area (SW Spain) in the wake of the 1998 mining waste spill. Mutagenesis 19: 61-65. doi: 10.1093/mutage/geg035

Jovani, R., & Serrano, D. (2004) Fine-tuned distribution of feather mites (Astigmata) on the wing of birds: the case of blackcaps Sylvia atricapilla. Journal of Avian Biology 35: 16-20. doi: 10.1111/j.0908-8857.2004.03213.x

Sol, D., Jovani, R., & Torres, J. (2003) Parasite mediated mortality and host immune response explain age-related differences in blood parasitism in birds. Oecologia 135: 542-547. doi: 10.1007/s00442-003-1223-6

Jovani, R. 2003. Understanding parasite strategies. Trends in Parasitology 19(1): 15-16.

Jovani, R. 2002. Malaria transmission, sex ratio, and erythrocytes with two gametocytes. Trends in Parasitology 18(12): 537-539.

Jovani, R., Tella, J.L., Blanco, G. & Bertellotti, M. 2002. Absence of haematozoa on colonial White Storks Ciconia ciconia throughout their distribution range in Spain. Ornis Fennica 79: 41-44.

Figuerola, J. & Jovani, R. 2001. Ecological correlates in the evolution of moult strategies in Western Palearctic passerines. Evolutionary Ecology 15:183-192. doi: 10.1023/A:1014824700389

Jovani, R. & Serrano, D. 2001. Feather mites (Astigmata) avoid moulting wing feathers of passerine birds. Animal Behaviour 62: 723-727. doi:10.1006/anbe.2001.1814

Jovani, R., Tella, J.L., Forero, M.G., Bertellotti, M., Blanco, G., Ceballos, O. & Donázar, J.A. 2001. Apparent absence of blood parasites in the Patagonian seabird community: is it related to the marine environment? Waterbirds 24: 430-433.

Jovani, R., Tella, J.L, Sol, D. & Ventura, D. 2001. Are hippoboscid flies a major mode of transmission of feather mites?. Journal of Parasitology 87: 1187-1189. doi:;2

Figuerola, J., Jovani, R. & Sol, D. 2001. Age-related habitat segregation by Robins (Erithacus rubecula) during the winter. Bird Study 48: 252-255. doi: 10.1080/00063650109461225

Jovani, R. & Blanco, G. 2000. Resemblance within flocks and individual differences in feather mite abundance on long-tailed tits Aegithalos caudatus. Écoscience 7(4): 428-432.

Tella, J.L. & Jovani, R. 2000. Sources of variability in aggregation and sex ratios of Crataerina melbae (Diptera:Hippoboscidae) among adult colonial alpine swifts. Journal of Parasitology 86: 933-938.

Sol, D., Jovani, R. & Torres, J. 2000. Geographical variation in blood parasites in feral pigeons: the role of vectors. Ecography 23: 307-314. doi: 10.1111/j.1600-0587.2000.tb00286.x